Evolution
Must Cooperation trigger, and what happens when the neighbour is already full or the Watering Hole is empty?
The designer confirmed Cooperation fires only on actual food taken and that a species that cannot feed passes nothing on.
Cooperation is mandatory whenever the species takes food. The neighbour receives food only if it can take it, so a species already at full food passes nothing on, while a Fat Tissue species that is still hungry can receive it. Nothing is taken from an empty Watering Hole, so Cooperation does not fire.