On Mars
How does the Remaining Missions marker move and trigger the end?
It moves one space each time a Mission is completed and each time the colony reaches a new level (3 or 4), even if a Mission is already done, so level-ups matter. Levels 1 and 2 do not move it. The end is triggered when the marker reaches the end of its track. In the printed rules every level-up advances it; a fan-made Tabletop Simulator board that disagrees is outdated.